Built to contract 11804 for the Ministry of Supply, dated February 1955. Officially on Ministry of Supply charge from 9 November 1958 and delivered via Filton on 12 November 1958. G-APPE carried spare parts for G-ANBC to Rangoon in record time on 13 March 1959, operating in civil markings. Royal Air Force Transport Command colours were applied in late March 1959, serialled XN392 she was named 'Acrux'. Evaluated by A&AEE Boscombe Down until 24 May 1959, she was damaged by salt spray at Belfast, causing her return to Filton for repairs to underside of fuselage and wings. Delivered to RAFTC No.99 Squadron at RAF Lyneham on 27 October 1959 she was operated within Nos. 99 and 511 Squadrons' Britannia pool at RAF Lyneham from 29 March 1961.

Withdrawn from service and stored at Baginton, Coventry from December 1975, XN392 had flown 12 652 hours and made 10 380 landings whilst in service with the RAF by the end of December 1975. Purchased by Aer Turas Teorante on 18 December 1975 she was broken up and scrapped during April and May 1976 after operation by City Airways fell through.