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Marines

The commanding officer of the ship's Marine detachment,, occupies a somewhat similar position with respect to the administration of the Marines aboard in matters pertaining strictly to the Marine Corps.

The role of the marine complement also varies a great deal from ship to ship.
The marine commander is accountable to the CO for the discipline of his marines

While the security department is responsible for the discipline of the crew, the marines (when they are present on board a ship) are responsible for the ship's safety--garrisoning sensitive areas, repelling boarders, etc. and for protection of away team missions. They also handle dignitary details, provide color guards for ceremonies, etc.  When marines are providing such services, they report to the XO. 
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