Lipid Structure and Metabolism

George M. Helmkamp, Jr., Ph.D.


Lecture 1: Lipid Structures; Digestion and Absorption (Powerpoint)

Learning Objectives

Lecture 2: Lipids as Sources of Metabolic Energy (Powerpoint)

Learning Objectives

Lecture 3: Synthesis of Structural and Bioactive Fatty Acid Derivatives (Powerpoint)

Learning Objectives

Lecture 4: Synthesis of Structural and Bioactive Polyisoprenes (Powerpoint)

Learning Objectives

Lecture 5: Medical Challenges of Cellular Lipid Metabolism (Powerpoint)

Learning Objectives)

Lecture 6: Vascular Lipid Metabolism: Overview and Dyslipidemias (Powerpoint)

Learning Objectives)

Lecture 7: Vascular Lipid Metabolism: Obesity and Diabetes (Powerpoint)

Learning Objectives)

Lecture 8: Vascular Lipid Metabolism: Atherosclerosis (Powerpoint)

Learning Objectives)


To TopLecture 1: Lipid Structures; Digestion and Absorption

Understand the nomenclature for fatty acids and complex lipids.

Know general structural features and functions of glycero-and sphingolipids.

List pancreatic enzymes involved in the digestion of dietary lipids.

Outline the formation of chylomicrons in the small intestine.

Explain why triacylglycerols are the preferred storage form of metabolic energy in humans; compare the synthesis of triacylglycerols in liver, adipose, and small intestine.

Describe the mobilization of adipose triacylglycerols and its hormonal regulation.

 

To TopLecture 2: Lipids as Sources of Metabolic Energy

Describe the activation of fatty acids to fatty acyl-coenzyme A thioesters.

Explain how fatty acids utilize carnitine to gain access to the mitochondrial matrix.

Outline the steps in the cycle of beta-oxidation of long-chain, saturated fatty acids; know the general chemical modifications to the acyl-CoA molecule.

Compare the plasma transport of fatty acids and ketone bodies and their use as metabolic fuels.

Be able to calculate the energy yield (ATP equivalents) from the oxidation of a saturated fatty acid to CO2 and H2O.

 

To TopLecture 3: Synthesis of Structural and Bioactive Fatty Acid Derivatives

Describe the function and organization of acetyl-CoA carboxylase and fatty acid synthase.

Outline the synthesis of long-chain, saturated fatty acids from acetyl-CoA.

Know sources of NADPH that are available for fatty acid synthesis.

Describe pathways of fatty acid elongation and desaturation; explain why linoleate and linolenate are essential to the human diet.

Compare the regulation and subcellular localization of fatty acid oxidation and synthesis.

Outline the synthesis of phosphatidate and diacylglycerol from glycerol 3-phosphate and fatty acids.

Outline the synthesis of ceramide from serine and fatty acids.

 

To TopLecture 4: Synthesis of Structural and Bioactive Polyisoprenes

Know the numbering system for the carbon atoms of cholesterol.

List major stages in cholesterol synthesis; note important building blocks and intermediates.

Discuss the regulation of 3-hydroxy-3-methylglutaryl-CoA reductase.

Outline cytochrome P-450-catalyzed hydroxylations of the sterol nucleus and side chain; describe the synthesis of pregnenolone as an important initial step in steroid hormone synthesis.

Describe general structures of bile acids and bile salts; explain their role in the emulsification and absorption of dietary lipids.

Recognize other biologically important polyisoprenes.

 

To TopLecture 5: Medical Challenges of Cellular Lipid Metabolism

Explain how phospholipases and other enzymes can re-model the structure of a complex lipid.

Describe the formation and function of pulmonary surfactant; summarize the characteristics of respiratory distress syndrome and its treatment.

Discuss general aspects of impaired sphingolipid degradation and lysosomal storage diseases; summarize the defect in Gaucher's disease and current treatment strategies.

Know some specific examples of the participation of the peroxisome in lipid metabolism; summarize the defect in Refsum's disease.

 

To TopLecture 6: Vascular Lipid Metabolism: Overview and Dyslipidemias

List the classes of plasma lipoproteins and describe their lipid transport functions.

Describe the enzymes involved in lipoprotein metabolism.

Discuss the metabolism of plasma lipoproteins and their specific interactions with cell-surface receptors; distinguish among the apo-B,E, apo-E, and scavenger receptors.

Define the process of reverse cholesterol transport; describe the involvement of high density lipoproteins in the transport and metabolism of cholesterol.

Summarize the clinical consequences of hypo- and hyperlipoproteinemias.

Understand the therapeutic value of medium chain triacylglycerols.

Describe the role of apo E in lipid metabolism; know the specific involvement of specific isoforms in vascular and central nervous system diseases.

 

To TopLecture 7: Vascular Lipid Metabolism: Obesity and Diabetes

List some causes (genetics and others) of human obesity.

Describe the role of leptin as a regulator of food intake, energy expenditure, and adipose tissue mass.

Discuss the hormonal regulation of adipose triacylglycerol synthesis and hydrolysis.

Explain why excessive plasma ketone bodies develop in poorly controlled type 1 diabetes mellitus.

Explain why elevated plasma triacylglycerols are characteristic of type 2 diabetes mellitus.

 

To TopLecture 8: Vascular Lipid Metabolism: Atherosclerosis

List risk factors associated with cardiovascular disease; distinguish between nonmodifiable and potentially controllable factors.

Describe the structure and development of atherosclerotic lesions; list the cells that participate in their formation.

Outline the consequences of impaired metabolism of low density lipoprotein in familial hypercholesterolemia.

Discuss the roles of modified LDL, lipoprotein(a), and homocysteine in cardiovascular disease.

Understand current therapeutic approaches to the treatment of elevated plasma lipids.

 

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