Homo sapiens neanderthalensis, 1856 Carolus Linnaeus
Neanderthal skulls were discovered in Engis, Belgium, in 1829 and Forbes' Quarry, Gibraltar, in 1848 prior to the "original" discovery in a limestone quarry of the Neander Valley (near Düsseldorf) in August, 1856, three years before Charles Darwin's On the Origin of Species was published.
H. Neanderthalensis, evolved from H. heidelbergensis in Europe about 300,000 y. a. (oldest find at Sima de los Huesos, Atapuerca)
Though Darwin's first book on evolution did not address the specific question of human evolution— "light will be thrown on the origin of man and his history," was all Darwin wrote on the subject— the implications of evolutionary theory were clear to contemporary readers. Debates between Thomas Huxley and Richard Owen focused on the idea of human evolution. Huxley convincingly illustrated many of the similarities and differences between humans and apes in his 1863 book Evidence as to Man's Place in Nature. By the time Darwin published his own book on the subject, Descent of Man, it was already a well-known interpretation of his theory— and the interpretation which made the theory highly controversial. Even many of Darwin's original supporters (such as Alfred Russel Wallace and Charles Lyell) balked at the idea that human beings could have evolved their apparently boundless mental capacities and moral sensibilities through natural selection.
Since the time of Carolus Linnaeus, the great apes were considered the closest relatives of human beings, based on morphological similarity. In the 19th century, it was speculated that our closest living relatives were chimpanzees and gorillas, and based on the natural range of these creatures, it was surmised humans share a common ancestor with African apes and that fossils of these ancestors would ultimately be found in Africa.
