One fan will run when the switch in the radiator reaches it's set point, and the ignition is off. Both fans will run when the switch in the radiator reaches it's set point, and the ignition is on, or when the A/C and the ignition are on. What happens is the radiator switch either closes a path through the resistor (with the ignition off), or closes a path through the cooling fan relay (with the ignition on). The cooling fan relay is also energized by the A/C relay. When the cooling fan relay operates, it bypasses the resistor and applies 12 volts to both fans. Also, the A/C relay inhibits the fresh air blower.