Cells as Experimental Models

E. coli , Yeasts , Dictyostelium discoideum , Caenorhabditis elegans, Drosophilia melanogaster

Arabidopsis thaliana , Vertebrates

Tools of Cell Biology: Light Microscopy, Electron Microscopy

Subcellular Fractionation

Growth of Animal Cells in Culture, Culture of Plant Cells

Viruses

Origin and Evolution of Cells:

Both Prokaryotes and Eukaryotes originated from a common single ancestor. First Cell is believed to have arisen as a result of enclosure of self-replicating RNA by phospholipid membrane about 3.5 to 3.8 billion years ago.

Stanley Miller demonstrated in 1950 the spontaneous formation of organic molecules by subjecting electric discharge to a mixture of H2, CH4, NH3 and H2O.

Evolution of Metabolism: Earliest organism obtained metabolic energy through a form of anaerobic glycolysis.

ATP generations are thought to have evolved in three stages; corresponding to the evolution of glycolysis, photosynthesis, and oxidative metabolism.

Prokaryotes have two division: Archaebacteria-which are progenitors of the most primitive bacteria and Eubacteria-modern day bacteria

Eukayotic Cells: Larger and more complex than Prokaryotes with its nucleus, cytoplasmic organelles and cytoskeleton, which are believed to have evolved from symbiotic association of prokaryotes (Endosymbiosis).

 

 

Cytoplasmic organelles:

Mitochondria and Chloroplast- Play very critical role in energy metabolism. Both organelles have DNA. Mitochondria is the site of oxidative metabolism and responsible for generating most of the cellular ATP.

Chlroplast- Site of Photosynthesis and found only in green plants and algae.

Lysosome and Peroxosome: Specialized metabolic compartment for digestion of macromolecules and many oxidative reactions.

Peroxisome- Involved in metabolism of H2O2, lipids and carbohydrate.

Vacuole- Digestion of macromolecules, storage of waste products, mostly found in plants.

Endomembrane System:

Rough endoplasmic raticulum-Synthesis and processing of protein.

Smooth Endoplasmic raticulum- Synthesis and processing of lipids, detoxification.

Golgi apparatus: Process, sort and transport protein, lipids and carbohydrates to their final destination. In plant cells serve as site for lipid synthesis and also polysaccharide synthesis for cell wall.

Multicellulor Organism:

Plants- Composed of fewer cell types compare to animal. Three main tissue system in plants-Parenchyma Cells- Carry out most of the metabolic reactions

Ground Tissue-Collenchyma and Sclerenchyma-Thick cell walls and provide structural support to plants.

Epidermal Cells- Protective covers and allows absorption of nutrients.

Animal-Human Body-200 different types of cells. Five different Tissue Types: Epithelial-Covers surface of body and line internal organs.

Connective Tissue-Bone, Cartilage, Adipose tissue formed by different types of cells such as Osteoblast, Chondrocytes, adipocytes, fibroblast.

Blood-Function in Oxygen Transport (Erythrocytes), Immune response (Granulocytes, Monocytes and Macro phages)

Cells as Experimental Model:

E.Coli- Genetic simplicity, easy to grow, regenerates and manipulates. Use for study of basic aspects of molecular biology and biochemistry.

Yeasts- Simplest eukaryote. Model for investigating eukaryotic cell biology,

Dictyostelium discoideum- Single cell eukaryote, use for studying cell movement.

Caenorhabditis elegans- Nematode, simple muticelluor organism useful as a model for developmental biology.

Drosophila melanogaster- Short generation time. Easy to manipulate, provided major clues in understanding animal development.

Vertebrates- Xenopus laevis-important model for studies of early vertebrate development.

Mouse- Mammalian species suitable for genetic analysis.

 Cell Culture:

Animal Cell: H. eagle 1955 first developed defines cell culture media.

Primary Cell Culture- Starting culture established from a tissue. Usually grow until culture dish surface is covered (Confluent Growth).

Immortal Cell Line- Cell culture derived from tumors proliferate indefinitely.

Plant Cell-Totipotency- cultured plant cell can differentiate to form specialized cell types, in some cases regenerate to produce whole plants.

Viruses:

Intracellular obligate parasites. Can be cultured only in living cells. Viruses consist of a nucleic acid core (DNA or RNA) and a protein cover.

Retroviruses- Family of RNA containing animal viruses containing reverse transcriptase which can synthesize a DNA copy of its genome.

Bacteriophage-Viruses that parasitize viruses.

 

 

 

 

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